Desaturase involved in fatty acid biosynthesis (PubMed:2118597). Introduces a double bond at carbon 12 of oleoyl groups (18:1) attached to the sn-1 position of the glycerol moiety of membrane glycerolipids (PubMed:20377689, PubMed:2118597). This enzyme is involved in chilling tolerance because the phase transition temperature of lipids of cellular membranes depends on the degree of unsaturation of fatty acids of the membrane lipids (PubMed:20377689, PubMed:2118597).